The rail-babbler and New Guinea species are birds of the rainforest, occupying a variety of closed forest types. The greater melampitta is restricted to forest on rugged karst environments. In Australia, only the eastern whipbird (P. oli-vaceus) is found in rainforest; it also occurs in wetter eucalypt forests and other low dense vegetation. The western whip-bird (P. nigrogularis) occurs in dense heath and other thick, drier vegetation. Wedgebills are birds of dry woodlands, steppes, and heathlands. Other than the spotted quail-thrush (C. punctatum), which prefers dry sclerophyll forest with open understorey, the other Australian quail-thrushes live in a range of arid habitats, including dry woodlands, shrub steppe, stony plains, and sandhills.

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